Signal charge that has accumulated in photodiodes of odd-numbered rows is read out one row at a time in turn in the column direction and represents an odd-numbered-row image. Signal charge that has accumulated in photodiodes of even-numbered rows is read out one column at a time in turn in the row direction and represents an even-numbered-row image. The even-numbered-row image is selected if a subject moves in the horizontal direction, and the odd-numbered-row image is selected if the subject moves in the vertical direction. An image in which rolling shutter distortion has been reduced is thus obtained.